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Cited article:

Increased complexity of gene structure and base composition in vertebrates

Ying Wu, Huizhong Yuan, Shengjun Tan, Jian-Qun Chen, Dacheng Tian and Haiwang Yang
Journal of Genetics and Genomics 38 (7) 297 (2011)
https://doi.org/10.1016/j.jgg.2011.06.004