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Cited article:

Fertilization and fertility in the female of Varroa destructor, a key point for the parasite population dynamics

Sébastien Wendling, Bertrand Guillet, Lise Roy, Serge Kreiter and Marc-Edouard Colin
Apidologie 45 (6) 722 (2014)
https://doi.org/10.1007/s13592-014-0291-4